2012–2015 Univ. Tokyo, Japan | Ph. D student
2015–2018 National Institute for Basic Biology, Japan | Postdoctoral fellow
2018–2020 Osaka Pref. Univ., Japan | Postdoctoral fellow
2020–2022 Tohoku Univ., Japan | Assistant professore
2022– CAS CEMPS/ CAS-JIC CEPAMS | Group leader
